I suspect there's another paper in the pipeline ... this one has no mention of the genetic relatedness of feral and managed colonies, but a few large hints in the intro and discussion. There's an interesting difference between the prevalence of DWV in the colonies reported here and the recent study by Furst et al., ... 100% here, 34% (if my increasingly dodgy memories serves me correctly) in the Nature paper.

Kate shows similar levels of DWV in untreated managed colonies and feral colonies. We know what happens to the majority of the former in a bad winter ...