Funnily enough my intitial searches all return anecdotal accounts on Scottish Beekeeping Association Sites.
There's a study from way back in 1989, but I've only just noticed that the springer.com links want to charge €34 to read the full report. More digging to follow...
Nuts, I wrote loads of notes around this point as, knowing I'd treated my colonies a couple of weeks apart, it was a point that pretty much made me sit bolt upright. I just don't think he mentioned the name of the actual study that was being carried out at the time or the one(presumably) that they used to get these figures.
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