Didn't Ian Rumsey (RIP) with his 'Pure and Simple' system find that the bees tended to align combs (given a free choice in a blank box) more according the orientation in their last home rather than anything else? DR, does your Apidea shelf face away from the main direction of your hives by 90 degrees?
Actually you might be right about that Gavin because the hive I stocked those mininucs from faced South
The offending mininucs were on a shelf in light shade facing North
Smith hives are not square and the entrance is (normally) on the long side
So those combs would be running North to South
The mininuc frames would be running East West
You might have solved that puzzle
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