No, nectar doesn't normally go from the crop to the ventriculus, at least not in quantity. The proventriculus sticks out into the crop or honey stomach. It manages to filter particles, gather them up, and pass them in a lump down into the ventriculus without there being a significant flow of nectar in that direction. The nectar remaining in the crop gets depleted of particles.
Which was:
(A fascinating feature of the proventriculus is that its four lips sieve out pollen, and even some bacteria and spores, allowing filtered nectar to pass through into the ventriculus.)
*Some* might make that jouney but most doesn't. The filtered nectar stays in the crop until the bee is ready to regurgitate, the solid matter heads down the digester. So the first sentence is right about the 4 lips filtering out particles, but wrong that a passage of nectar to the ventriculus is involved.
Do have a go at the Bailey paper mentioned above. Access to the full PDF is free.
Bookmarks